Each of the checklists
is based on a single taxonomic source that covers all species in a
group on a global basis. For the sake of consistency, each of
these taxonomic sources is used as the final authority on the validity
of species regardless of subsequent taxonomic changes or new
discoveries. Although most of these sources also included some
information on distribution, all required checking additional sources
to confirm ranges. Islands or island groups divided into more
than one political unit (such as Borneo, Hispaniola, the Solomons, and
the Comoros) almost invariably required such checking.
An endemic species for the purposes of the checklists is considered to
be one that currently occurs as a breeding species in a single
political unit (for example New
Caledonia or South Sudan).
Formerly more widespread species that
currently survive in only a single country are also considered endemic
(for example the Hirola in Kenya).
Species that have been introduced to other countries are also
considered endemic to their original country of origin (for example South Africa's Bird
of Paradise Flower).
The
situation is more complex for the handful of species in
which reintroduction attempts are being made into their former range.
For example, the Crested Ibis was included in the checklist
as
endemic to China
until reintroduced birds in Japan
successfully bred in 2012.
Species known from border regions and frequently assumed to occur also in similar habitats in a neighboring nation are treated as endemic unless a definite record exists for the second country (an exception is freshwater fishes collected on a stretch of river that serves as a border which are assumed to occur in both countries even if specimens are known only from one bank).
In
cases of doubt species are usually included as an
endemic. Recently extinct species (such as the Golden Toad in Costa Rica) are included in the
lists
if they were included in the taxonomic source, both because they may be
indicators of areas that may include other endemic species and because
some "extinct" species have turned out to be not quite as extinct as
had
been previously assumed (for example: the Forest Owlet in India).
How
accurate are the checklists?
It is safest to assume that most of the longer checklists
(and many of the shorter ones) can be improved upon. Although
for some island nations the checklists may accurately reflect current
knowledge, for many mainland nations the situation is quite a bit
messier. In a number of cases a species will be included as
an endemic because an inadequate search of the literature by the
compiler failed to discover records from other countries.
One complicating factor is a lack of consensus on species boundaries that can result in misidentifications in the literature or differing usages of the same scientific name to mean different things. For example the name Antidorcas marsupialis is most commonly used to refer to an antelope found in several countries (IUCN), but has been used at times in a more narrow sense to refer to a population occuring only in South Africa (Google Books).
A number of species may have
been inadvertantly omitted from the list by the compiler not realizing
that a name was being used in a different sense. Misidentifications in
databases such as GBIF
are unfortunately all too frequent and one study (Current
Biology)
concludes that perhaps as many as half of all museum tropical plant
specimens
have
been misidentified creating additional problems in determining accurate
species ranges. Specimen data are subject to a variety of possible
location errors (BHL).
Museum databases also sometimes misinterpret collection localities to
an incorrect modern country (and sometimes continent).
It is also important to remember how much is still unknown to science.
Thousands of vertebrate species, tens of thousands of
vascular plant species, and millions of invertebrate species still
await discovery and scientific description. For example,
according to the State of Observed Species report 19,232
new species were described in 2009 (ESF
WM
pdf). Over 8000 new species of fish have been
described since 2004 (Catalog
of Fishes).
There is often also a long lag time between the first collection of a
species and its scientific description. One recent study found that
this averages over 21 years (ResearchGate).
Distributions for many species, especially in the tropics, are also incompletely known with much of the world very poorly sampled (NSO pdf). For example, one in seven lizard species have been found solely in the location where it was originally discovered (ResearchGate).
Some
parts of the world are also much
more poorly
known than others - for example only two of the 105 freshwater fish
species known solely from Laos
had even been described before 1995.
A sizeable majority of the species newly described are known
from only a single nation. 205 out of 220 (93.2
%) reptile species described in 2019 (according to Reptile
Database accessed 25 May 2020) and 138 out of 149 (92.6
%) amphibian species described in 2019 (according to AmphibiaWeb
accessed 11 April 2020) are so far
known exclusively from a single country.
Political
Units
Countries and dependent areas are those recognized by the CIA World
Factbook (CIA).
Additional accounts appear for species of Unknown
Origin,
those found solely in International
Waters, those unique to the Korean
Peninsula, and those restricted to the island of Timor. The Cyprus account includes the
nation
of Cyprus plus the United Kingdom
administered areas of Akrotiri and Dhekelia.
Israel, the West Bank, and the
Gaza Strip are discussed
jointly for practical reasons. Threatened endemic species can be found
using the Advanced Search function at the IUCN Red List
which includes filters for countries and endemics (near the bottom of
the page).
Country
Overview Pages
These pages provide introductions to the wealth of resources on the web
dealing with the world's animal and plant species. It also
allows for the mention of endemic invertebrate and vascular plant
species plus additional vertebrate species that are recognized by some
authorities or were described since the sources used for the
checklists. Countries and dependent areas lacking known
endemic
species are not given a country overview page.
Mammals
American Society of Mammalogists (2020) Mammal Diversity Database,
Version 1.2 https://www.mammaldiversity.org,
accessed 24 September 2020.
An online version of the above work can be found here.
Lists of additional species recognized by the Illustrated Checklist of the Mammals of the World (Lynx) are also included. Mammal species unique to a single nation added to the Mammal Diversity Database after this date are periodically included as addenda in the appropriate national checklists.
The MDD recognized 6485 recent mammal species of which 2896
species (44.7 %) are single nation endemics. 307 of 1330
(23.1 %) mammal genera are also restricted to a single country.
Birds
del Hoyo, J. & N.J. Collar. HBW
and BirdLife International Illustrated Checklist of the Birds of the
World.
2014 Vol. 1: Non-passerines & 2016 Vol. 2: Passerines. Lynx
Edicions, Barcelona. The lists are based on the December 2017 update
available
here.
11,122 bird species are recognized in the December 2017 update of which
3450 (31.0 %) are
single nation endemics. Bird species unique to a single nation
recognized after the December 2017 update are periodically
included
as addenda in
the appropriate national checklists. Another addenda indicates
additional species recognized as valid by one of the following
checklists: IOC 10.1 (IOC),
Howard & Moore 4.1 (TAS),
and eBird/Clements v2019 (Cornell).
367 of 2365 (15.5 %)
bird genera are
also restricted to a single country. An independent
assessment of single nation endemics using the above reference can be
found at Avibase (Avibase)
which also provides assessments based on several other authorities as
well. See also (Lynx pdf).
An especially detailed source of distributional
information for all the world's birds can be found at BirdLife
International (BirdLife
Int'l). For recently described bird species see (listaptakow.eko.uj.edu.pl).
Reptiles
Uetz, P.,
Freed, P. & Hošek, J. (eds.) (2018), The Reptile
Database, https://www.reptile-database.org, accessed 14
November 2018.
The current online version of the
Reptile Database can be found here.
The 14
November 2018 version of the Reptile Database recognized
10,885
reptiles species of which 7060 (64.9 %) are single nation endemics.
Reptile species unique to a single nation added
to the Reptile Database after this date are periodically included
as addenda in
the appropriate national checklists. 373
of 1193 (31.3 %)
reptile genera are also restricted to a
single country. For lists of very recently described reptile
species see (Reptile
Database) (Reptile Database) or search by year and location at (Reptile
Database).
Amphibians
AmphibiaWeb (based
on species recognized as of 28 October 2018) The current online
version of this database can be found here.
The 28 October 2018 version of AmphibiaWeb recognized 7941 species of which 5816 (73.2 %) are single nation endemics. Amphibian species unique to a single nation added to AmphibiaWeb after this date are periodically included as addenda in the appropriate national checklists. 179 out of 547 (32.7 %) amphibian genera are also restricted to a single country. For lists of recently described amphibian species search by year of publication and country at (Amphibian Species of the World) or see the new species page at AmphibiaWeb (AmphibiaWeb). Lists of endemic species by country can be generated at (Amphibian Species of the World).
Fricke, R., Eschmeyer, W. N. & Van der Laan, R. (eds) 2023 Catalog of Fishes: Genera,
Species, References. Version downloaded 9 February 2023. The current version can be
accessed here
18,535 species of fish are found in freshwater or brackish habitats
(including marine species extending into freshwaters, species only
known from brackish waters, but not incuding marine species that extend
into brackish waters but not freshwater) according to Fricke, et
al. 2023. 10,914 (58.9 %) of these species
are single
nation endemics.
The marine fish checklists include only species that occur mainly in habitats between 0 and 200 m deep and therefore usually do not include bathypelagic and bathydemersal species. Additional lists of deep-sea species (below 200 m) known from a single country are listed at the bottom of each page. Maritime boundaries follow those mapped at (marineregions.org).
About 932 out of 5229 (17.8 %) of fish genera are restricted to a single country. About 15,922 fishes (including deep-sea fishes) are known from a single country out of 36,415 total fish species known (43.7 %) according to Fricke, et al. 2023..
For
recently described fish species
see (worldfish.de)
and search by year and location at (Catalog
of Fishes) or by year at (FishBase).
Vertebrate
Genera
The sources for the vertebrate genera are the same as those for
species. However some genera have been excluded as being
endemic based on the ranges of subsequently described or undescribed
species. Approximately 2158 out of 10,664 (20.2 %) vertebrate genera
are
single nation endemics.
Swallowtail
& Nymphalid Butterflies
Jonko, C.
(2019), Lepidoptera
Mundi, https://lepidoptera.eu, accessed 12 January
2020. The current version can be accessed here
These two butterfly families have been selected as examples of
invertebrate
endemism. 2030 out of 6691 (30.3
%) of these two butterfly families are single nation endemics.
National checklists at the species level are
available for a
number of other invertebrate groups through the 2019 Annual Checklist
of the Catalogue of Life using
the search by distribution feature (Catalogue
of Life).
Vascular
Plant Genera
Mabberley,
D.J. (2017) Mabberley's
Plant-book. Fourth Edition. Cambridge University Press,
Cambridge.
Mabberley recognized 13,763 vascular plant genera of which 3300 (24.0 %)
are single nation endemics. According to the
GlobalTreeSearch database (BGCI
pdf),
33,760 out of the world's 58,497 (57.7 %) tree species are single
country endemics.
Trees are believed to represent about 20 % of all angiosperm and
gymnosperm species. Endemic plant species are
indicated for most
countries in checklists that can be accessed at (worldplants.de). Note
that
it is also possible to
create endemic species checklists for
many plant families and countries using the "Build a Checklist" feature
at the World
Checklist of Selected Plant Families website (Kew).
Recently described species can be found using the advanced search
function at
the International Plant Names Index (IPNI).
Families
& Orders
The
sources for the vertebrates and vascular plants are the same as given
above.
For invertebrates the primary sources are Zhang (2013) and Zhang (2011). Roskov et al. (2014) is used as the main source for groups not covered at the family level by Zhang (2013) and Zhang (2011), but this was supplemented by other sources at times as it also has some gaps (for example with terrestrial gastropods). A number of more recently described families have also been included (usually indicated by year of description).
Out of 7177 families (1182 vertebrates, 5521 invertebrates, and 474 vascular plants), 512 (7.1 %) are single nation endemics.
Zhang, Z.-Q. (Ed.) (2013) Animal Biodiversity: An Outline of Higher-level Classification and Survey of Taxonomic Richness (Addenda 2013). Zootaxa 3703: 1–82. (here)
Zhang, Z.-Q. (Ed.) (2011) Biodiversity: An Outline of Higher-level Classification and Survey of Taxonomic Richness. Zootaxa 3148: 1–237. (here)
Roskov Y., Kunze T., Orrell T., Abucay L., Paglinawan L., Culham A., Bailly N., Kirk P., Bourgoin T., Baillargeon G., Decock W., De Wever A., Didžiulis V., eds. (2014). Species 2000 & ITIS Catalogue of Life, 2014 Annual Checklist. Digital resource at www.catalogueoflife.org/annual-checklist/2014. Species 2000: Naturalis, Leiden, the Netherlands.